Criticisms On “The Origin Of Species” Informative Summary


This text, penned by Thomas Henry Huxley in 1864, examines various critiques of Charles Darwin’s groundbreaking work, “On the Origin of Species.” Huxley, a staunch defender of Darwin’s ideas, dissects arguments put forth by German anatomist Albert Kölliker and French scientist Pierre Flourens. Both critics raise concerns about the lack of transitional fossils, the absence of useful variations, and the implications of Darwin’s theory for the concept of progress.

Huxley adeptly addresses these concerns, demonstrating that Darwin’s theory doesn’t require a continuous progression towards perfection but rather focuses on the survival of the fittest. He highlights that natural selection, a key element of Darwin’s theory, is not a conscious process but a natural consequence of environmental pressures and the inherent variability of organisms. Furthermore, Huxley defends Darwin’s teleological stance, arguing that Darwin’s focus on the benefits of adaptation doesn’t contradict his broader rejection of traditional teleological arguments. He refutes Flourens’ critique of natural selection as a mere “personification” of nature, highlighting the demonstrable effects of environmental pressures on organisms, akin to the way wind shapes dunes.

Key Findings:

  • Natural Selection as a Non-conscious Force: Huxley clarifies that Darwin’s theory of natural selection operates without conscious intent, much like the way the wind shapes dunes or a frost selects for hardy plants.
  • Rejection of Progress: Huxley challenges the misconception that Darwin’s theory implies a constant, linear progression towards perfection, emphasizing that it allows for stagnation and even retrogression.
  • Teleology Reconsidered: Huxley explains that Darwin, while rejecting traditional teleological arguments, emphasizes the utility of features and adaptations, thereby reconciling the apparent conflict between teleological and morphological views.


  • Natural Selection and Environmental Pressure: The reader learns that natural selection is not a deliberate force but a natural consequence of environmental pressures. Organisms with traits that best suit their environment are more likely to survive and reproduce, passing on those advantageous traits to future generations.
  • Variation and Adaptation: The text elucidates the importance of variation within species. This inherent variability provides the raw material for natural selection, allowing organisms to adapt to changing environmental conditions and exploit new niches.
  • Lack of Progress in Evolution: The reader gains a more nuanced understanding of evolution, recognizing that it doesn’t always result in a steady march towards perfection. Evolution can lead to stasis, retrogression, or even extinction based on the interplay of environmental pressures and variations within populations.
  • Understanding the Limits of Teleology: The text sheds light on the limitations of traditional teleological arguments, which posit that organisms were designed with a specific purpose in mind. Darwin’s work challenges this perspective, emphasizing the role of natural selection in shaping organisms over time.

Historical Context:

This text was written during a period of intense debate surrounding Darwin’s ideas. Scientific circles were grappling with the implications of evolution and the challenge it posed to traditional beliefs about the origins of life. The publication of “Origin of Species” in 1859 sparked controversy, and Huxley’s defense of Darwin’s ideas reflects this turbulent intellectual landscape.


  • Variation is Essential for Evolution: All organisms within a species exhibit variations in traits, providing the raw material for natural selection.
  • Environmental Pressures Drive Selection: The environment exerts selective pressure on organisms, favoring those with traits that best suit the prevailing conditions.
  • Adaptation is the Result of Selection: Organisms adapt to their environments over time through the accumulation of advantageous traits selected for by environmental pressures.
  • Evolution Can Lead to Stagnation: A species may remain unchanged for long periods if it is well-adapted to its environment and there is no significant environmental change.
  • Evolution Can Lead to Retrogression: Species can become less complex or even extinct if they are unable to adapt to environmental changes.
  • Evolution Does Not Always Lead to Perfection: Evolution is not a goal-oriented process. Organisms are not inherently striving to become more perfect, and evolution can result in organisms that are less complex or well-adapted.
  • Darwin’s Ideas Faced Strong Criticism: Darwin’s theory of evolution faced strong resistance from both the scientific community and the general public, as it challenged traditional beliefs about the origins of life.
  • Huxley Was a Vocal Advocate for Darwin: Thomas Henry Huxley was a staunch defender of Darwin’s theory and played a significant role in promoting its acceptance.
  • Kölliker Offered an Alternative Theory: Albert Kölliker, a prominent German anatomist, proposed an alternative theory to Darwin’s, based on the concept of “heterogeneous generation.”
  • Flourens Dismissed Darwin’s Work as Unsound: Pierre Flourens, a French scientist, vehemently criticized Darwin’s theory, dismissing it as metaphysical jargon and a personification of nature.
  • Agamogenesis Is a Different Phenomenon: The phenomenon of agamogenesis (asexual reproduction) cannot be used to explain the origin of new species, as it is a cyclical process that returns to the original type.
  • Variations Can Be Substantial: Variations within species can be significant, leading to substantial differences in traits.
  • Evolutionary Jumps Are Possible: Evolutionary change is not always gradual, and substantial variations can occur, potentially leading to the emergence of new species.
  • The Fossil Record Is Incomplete: The fossil record provides important evidence for evolution, but it is incomplete, making it difficult to document every transitional form.
  • Natural Selection Is a Powerful Force: Natural selection is a powerful force that shapes the diversity of life on Earth, leading to the adaptation of organisms to their environments.
  • Darwin Emphasized the Utility of Features: Darwin’s work focused on the usefulness of features, recognizing that they are products of natural selection and contribute to the organism’s survival.
  • Darwin’s Work Was Groundbreaking: Darwin’s theory of evolution revolutionized our understanding of the origins of life, setting the stage for modern evolutionary biology.
  • Darwin’s Ideas Faced Opposition from Traditional Beliefs: Darwin’s theory challenged the prevailing view of a divinely created and unchanging world, leading to widespread resistance and controversy.
  • Huxley’s Defense of Darwin Was Important: Huxley’s defense of Darwin’s theory was important for promoting its acceptance and helping to lay the foundation for modern evolutionary biology.


  • Darwin’s “Origin of Species” was published in 1859. This landmark work sparked a scientific revolution and continues to shape our understanding of the natural world.
  • Huxley’s essay on criticisms of Darwin’s work was published in 1864. This text provides valuable insight into the debates surrounding Darwin’s theory during a period of intense intellectual scrutiny.
  • Darwin’s “Origin of Species” has been translated into over 40 languages. This wide dissemination underscores its global impact and the importance of its ideas.
  • Kölliker’s critique of Darwin’s theory was published in 1864. This essay reflects the ongoing scientific debate surrounding Darwin’s ideas.
  • Flourens’ critique of Darwin’s work was also published in 1864. This text highlights the continued opposition to Darwin’s theory, even from prominent scientists.


  • Teleology: The philosophical doctrine that everything in the universe has a purpose or design. Traditional teleological arguments suggested that organisms were created with a specific purpose in mind. Darwin’s theory of evolution challenged this view.
  • Natural Selection: The process by which organisms with traits that best suit their environment are more likely to survive and reproduce, passing on those advantageous traits to future generations.
  • Agamogenesis: A form of asexual reproduction, also known as parthenogenesis, in which an organism develops from an unfertilized egg or ovum.
  • Variation: Differences in traits among individuals within a species, providing the raw material for natural selection.
  • Adaptation: The process by which organisms evolve traits that increase their chances of survival and reproduction in their environment.
  • Heterogeneous Generation: A theory proposed by Kölliker, suggesting that new species arise from pre-existing ones through a process of spontaneous generation.
  • Entwickelungsgeschichte: A German term for “evolutionary history.”
  • Foetal Balaena: A reference to the baleen whale, highlighting its teeth in the foetal stage, which disappear in the adult stage. This is an example of a feature that may have been useful to an ancestral form but no longer serves a direct purpose.
  • Ancon Ram: A well-known example of a significant variation, a breed of sheep with short, crooked legs, that arose spontaneously.


  • The Ancon Ram: This breed of sheep, with short, crooked legs, arose spontaneously from an ordinary ewe’s ovum. It is a compelling example of substantial variation that can occur within a species.
  • The Cats and Mice Example: Huxley uses the example of cats catching mice to illustrate the difference between teleological and Darwinian viewpoints. While teleology would argue that cats were designed specifically to catch mice, Darwinism suggests that cats exist because they are good at catching mice, and this ability has been selected for over time.
  • The Dunes of the Landes: Huxley uses the formation of dunes on the coast of France as an analogy for natural selection. The wind, without consciousness, selects for sand grains of a specific size, just as natural conditions select for organisms with advantageous traits.
  • The Thistle on the Pampas: The spread of the thistle across the pampas, displacing native plants, exemplifies the power of natural selection. The thistle’s ability to thrive in the pampas environment has allowed it to outcompete other plants, demonstrating the effectiveness of natural selection in shaping ecosystems.
  • Foetal Teeth in Baleen Whales: The presence of teeth in the foetal stage of baleen whales, which disappear in the adult stage, exemplifies a feature that may have been useful to an ancestral form but no longer serves a direct purpose.


Thomas Huxley’s essay effectively defends Darwin’s “Origin of Species” from the criticisms of Kölliker and Flourens. It clarifies key concepts such as natural selection, adaptation, and the role of variation in evolution. Huxley demonstrates that Darwin’s theory doesn’t necessitate a continuous progress towards perfection and that natural selection operates as a non-conscious force, driven by environmental pressures. By clarifying these points and addressing common misconceptions, Huxley underscores the validity of Darwin’s groundbreaking work and its significance for understanding the diversity of life on Earth.

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