The Darwinian Hypothesis Informative Summary


This essay, published in 1850 in the Times newspaper, is a response to Charles Darwin’s revolutionary work on the origin of species. Huxley argues that Darwin’s theory of natural selection presents a compelling and scientifically sound explanation for the diversity of life on Earth. He begins by exploring the challenges of defining species, noting the inconsistencies in how naturalists classify organisms and the difficulties in applying a strict definition of a species based on reproductive fertility.

Huxley then outlines the evidence supporting the idea of species change over time, drawing on geological evidence of fossils and the gradual evolution of life forms. He contrasts Darwin’s theory with Lamarck’s earlier hypothesis of evolution driven by acquired characteristics, highlighting the strength of Darwin’s concept of natural selection as a driving force for change. Huxley concludes by arguing that Darwin’s theory provides a more comprehensive and scientifically sound explanation for the patterns observed in nature than the traditional view of species immutability.

Key Findings:

  • Species are not fixed: The essay challenges the prevailing view that species are immutable, arguing that species change over time in response to environmental pressures.
  • Natural selection as a driver of change: Huxley highlights the role of natural selection in shaping the diversity of life, emphasizing how competition for resources and survival pressures lead to the adaptation and evolution of species.
  • The limitations of final causes: The essay criticizes the traditional reliance on teleological explanations for biological phenomena, arguing that Darwin’s theory provides a more scientific and testable framework for understanding the origin of species.


  • The concept of natural selection: Readers will learn about Darwin’s groundbreaking theory of natural selection, understanding how it explains the evolution of species through variations, inheritance, and competition for survival.
  • The evidence for evolution: Huxley presents various pieces of evidence that support the idea of evolution, including geological evidence of fossils, the distribution of species, and the development of organisms.
  • The limits of traditional explanations: Readers will learn about the limitations of traditional explanations for the diversity of life, such as the reliance on fixed species and teleological arguments.
  • The importance of scientific inquiry: The essay emphasizes the importance of scientific inquiry and the need to test hypotheses through observation and experiment.

Historical Context:

This essay was written in 1850, a time when Darwin’s theory of evolution was still largely unknown to the public. The traditional view of species being created by God was still widely accepted. This context highlights the significance of Huxley’s essay as one of the first public responses to Darwin’s work.


  1. The number of species is vast: There are at least 100,000 known species of insects and likely over half a million species of living things in total. This immense diversity reflects the complex processes of evolution.
  2. Species can hybridize: Contrary to the notion of species purity, some hybrids are fertile, showing that strict barriers between species are not absolute.
  3. Rudimentary organs exist: The presence of rudimentary organs, like vestigial teeth in whales or wings in flightless insects, supports the idea of evolutionary change and the loss of function over time.
  4. Animals and plants share early developmental stages: All living beings, from humans to sponges, begin their development in similar forms, highlighting common ancestry.
  5. Geographical distribution is complex: The distribution of species cannot be explained solely by adaptation to local environments, suggesting historical factors and the influence of migration and isolation.
  6. Fossils reveal past life: The vast fossil record provides a glimpse into the history of life on Earth, showcasing extinct species and the evolution of lineages over time.
  7. Extinct species are not simply replaced by similar ones: The fossil record shows a gradual change in life forms, with younger species often sharing similarities with older ones, but not being identical replacements.
  8. Species replacement is gradual: The transition between species is not sudden or catastrophic but occurs through a gradual process of adaptation and change.
  9. Breeders create artificial variations: The breeding of domesticated animals and plants demonstrates the ability to select and enhance traits, providing a model for how natural selection can operate.
  10. The struggle for existence is a natural phenomenon: The competition for resources and survival drives natural selection, leading to the survival of the fittest and the elimination of less well-adapted individuals.
  11. Every species produces more offspring than can survive: This overproduction ensures that there is sufficient variation for natural selection to act upon.
  12. Variations arise randomly: New traits or variations appear spontaneously within a species, providing the raw material for natural selection to work with.
  13. Favorable variations are passed down: Individuals with variations that make them better suited to their environment are more likely to survive and reproduce, passing these beneficial traits to their offspring.
  14. Natural selection is a continuous process: The process of natural selection is ongoing, leading to gradual changes in species over generations.
  15. The environment is a selective force: Changes in the environment can favor different traits, leading to the evolution of new species.
  16. The fossil record reveals transitional forms: The discovery of fossils that exhibit characteristics of both ancestral and descendant species supports the idea of gradual evolutionary change.
  17. Geographical isolation can lead to speciation: The separation of populations can lead to different selective pressures, resulting in the development of distinct species.
  18. Natural selection is a non-teleological process: Natural selection does not work towards a predetermined goal, but rather results from the interaction between random variations and environmental pressures.
  19. Humans are subject to natural selection: The essay points to the potential for humans to be affected by natural selection, suggesting that certain traits may be favored in different environments.
  20. Evolution is a continuous process without a final endpoint: Natural selection is a dynamic process that continues to shape life on Earth, constantly producing new variations and adaptations.


  1. Half a million species: This is a conservative estimate of the number of distinct living species on Earth, highlighting the incredible diversity of life.
  2. Vast time scales: The geological record indicates that the Earth is billions of years old, providing ample time for evolution to occur.
  3. Overproduction of offspring: Every species produces more offspring than can survive, ensuring the availability of variation for natural selection.
  4. Millions of cattle in the Pampas: This illustrates how introduced species can thrive in new environments, suggesting that adaptation is not always about existing species being perfectly suited to their native locations.
  5. Hundreds of thousands of insect species: This reflects the immense diversity of insects and their adaptability to various ecological niches.
  6. A hundred thousand known insect species: This statistic highlights the vastness of the insect world and the challenges of classifying such a diverse group.
  7. One to a million young: This range reflects the vast differences in reproductive potential among species, demonstrating the importance of survival rates in driving population dynamics.
  8. Twenty years of research: Darwin’s research, spanning over two decades, reflects the dedication and meticulous observation required to understand complex biological processes.
  9. The Earth’s vast history: The geological record provides evidence of a long and complex history of life on Earth, offering insights into the processes of evolution.


  • Species: A group of organisms that can interbreed and produce fertile offspring.
  • Hybrid: The offspring of two different species.
  • Natural selection: The process by which organisms with traits better suited to their environment are more likely to survive and reproduce.
  • Variation: Differences in traits among individuals within a species.
  • Adaptation: A trait that allows an organism to survive and reproduce in a specific environment.
  • Extinction: The disappearance of a species.
  • Fossil: The preserved remains of a prehistoric organism.
  • Rudimentary organ: A vestigial structure that has lost its original function.
  • Geographical distribution: The patterns of where species are found around the world.
  • Transmutation theory: The theory that species can change over time into new species.


  1. The diversity of domesticated animals and plants: The differences between breeds of cattle, sheep, pigeons, and dogs illustrate the potential for human selection to create dramatic variations.
  2. The evolution of flightless birds: The existence of flightless birds, like the ostrich and emu, with vestigial wings, supports the idea of evolution and the loss of function through natural selection.
  3. The distribution of species on islands: The unique species found on islands, like those described by Sir Emerson Tennent in Ceylon, highlight the role of geographical isolation in speciation.
  4. The lack of cattle in the Pampas: The absence of cattle in the Pampas of South America before their introduction demonstrates that adaptation is not always about existing species being perfectly suited to their environments.
  5. The case of the heron’s neck: Lamarck’s example of the heron’s long neck, which he attributed to acquired characteristics, is used to highlight the flaws in his theory compared to the more scientifically sound explanation of natural selection.
  6. The struggle for existence in nature: The competition for resources among plants and animals, as well as the high rates of mortality, illustrate the natural forces that drive natural selection.
  7. The development of the human embryo: The similarities between the development of human embryos and other animals, like fish and birds, point to a shared evolutionary history.
  8. The fossil record of horses: The sequence of fossils showing the gradual evolution of horses from small, multi-toed ancestors to larger, single-toed horses provides strong evidence for evolutionary change over time.
  9. The geographical isolation of Darwin’s finches: The diversity of finches on the Galapagos Islands, each adapted to a different food source, illustrates the role of geographical isolation in driving speciation.
  10. The evolution of antibiotic resistance in bacteria: This example demonstrates the rapid evolution of organisms in response to changing environmental pressures.


Huxley’s essay stands as a powerful early defense of Darwin’s theory of natural selection. By challenging traditional views of species immutability and exploring the evidence for evolution, Huxley highlights the revolutionary implications of Darwin’s work. He emphasizes that Darwin’s theory provides a more scientific and testable framework for understanding the diversity of life on Earth than previous explanations based on divine creation or acquired characteristics. The essay ultimately serves as a call for further investigation and a recognition of the transformative power of scientific inquiry in challenging long-held beliefs.

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